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Ruminant Digestive Physiology
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Esophagus
The esophagus is responsible for transporting food from the mouth to the rumen and reticulum, and also enables the regurgitation of cud for rumination.
Saliva
Saliva in ruminants buffers the rumen pH and provides liquid for the microbial population, which is important for the fermentation process.
Lipid Metabolism
Lipids in the diet are hydrolyzed by rumen microbes into glycerol and free fatty acids, which undergo biohydrogenation to become more saturated.
Particle Size Reduction
Rumination and microbial digestion reduce the particle size of feed, facilitating better digestion and passage of the digesta through the rest of the digestive system.
Reticulum
The reticulum works closely with the rumen for fermentation but is also the site where foreign objects are trapped, minimizing potential damage to the digestive tract.
Rumen
The rumen is a large fermentation vat where microbial digestion of fibrous plant material occurs, producing volatile fatty acids which are a primary energy source for ruminants.
Rumination
Rumination is the process of chewing cud to mechanically break down fiber, which increases the surface area for microbial digestion in the rumen.
Omasum
The omasum absorbs water and nutrients from the digesta and also helps in reducing the particle size of the digesta.
Waste Excretion
The ruminant digestive system allows for the excretion of undigested waste and non-absorbable materials efficiently through feces, which also contributes to the animal's nutrient cycle.
Microbial Population
Microbial populations in the rumen synthesize protein from non-protein nitrogen sources, produce B vitamins, and aid in the detoxification of harmful compounds.
Eructation
Eructation is the release of gases, mainly carbon dioxide and methane, produced during microbial fermentation, preventing gas buildup and bloat in the animal.
Volatile Fatty Acids (VFAs)
VFAs like acetate, propionate, and butyrate are the primary products of fermentation in the rumen and are absorbed through the rumen wall, providing a significant energy source.
Abomasum
The abomasum is the true stomach where chemical digestion takes place using gastric juices to breakdown proteins and non-fibrous components of the feed.
Rumen Papillae
Rumen papillae increase the surface area for the absorption of volatile fatty acids and help protect the rumen wall from mechanical damage by feed particles.
Detoxification
Rumen microbes can detoxify various plant toxins and anti-nutritional factors before they enter the host animal's systemic circulation, enhancing food safety.
Rumen Motility
Rumen motility is crucial for mixing and moving the contents of the rumen, ensuring even fermentation and preventing stratification of the digesta.
Rumen Mat
The rumen mat is a layer of fibrous feed particles that float on top of the rumen liquid, aiding in mechanical digestion and regulating the passage of finer particles.
Carbohydrate Digestion
Carbohydrates from plant cell walls, like cellulose and hemicellulose, are digested by rumen microbes into simple sugars that can be fermented to VFAs.
Rumen Synthesis of Micronutrients
Micronutrients such as B vitamins and vitamin K are synthesized by the rumen microbes, reducing the dietary requirement for these vitamins.
Protein Digestion
In the rumen, dietary proteins are broken down by microbes into peptides, amino acids, and ammonia, which can be used by the microbes or absorbed as non-protein nitrogen.
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